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Oakmead Herbarium: Undocumented reports and erroneous inclusions

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Table 1. Undocumented reports.

These historical records do not meet the minimum criterion for inclusion on the Jasper Ridge plant list for unvouchered plants, namely, reported by more than one observer with detailed location information. For some species and genera (e.g., Arctostaphylos, Carex, Cryptantha, Lemna, Plectritis, Salix) we also required mention of key morphological characters. Some of the plants reported, in any case, are garden escapes and waifs not expected to naturalize. All Springer 1935 and Moeur 1947 reports lack vouchers. Furthermore, Moeur appears largely derivative of Springer's flora. Dengler 1975 and his other lists comprising 600+ reports are supported by 94 sheets for 85 taxa; some taxa are mentioned in his notebooks and elsewhere. Most of the plants below were only reported on Dengler's 1975 plant list.

APIACEAE

  • Apiastrum angustifolium — Dengler 1975; not reported since. No voucher; not expected habitat. See note
  • Perideridia gairdneri ssp. gairdneri — Moeur 1947. See note

AMARANTHACEAE

  • Amaranthus blitoides — Dengler 1975; not reported since

ASTERACEAE

  • Anaphalils margaritacea — Springer 1935 and Moeur 1947; not reported since. Probably Pseudognaphalium californicum
  • Arctium lappa — Dengler 1975; not reported since
  • Crepis bursifolia — Perennial, base of plant swollen and ± woody; taproot stout. Macklin 1988; not reported since. CCH records for Stanford Campus
  • Lasthenia glaberrima — Dengler 1975; not reported since
  • Packera breweri (Senecio b.) — Moeur 1947; not reported since

BORAGINACEA

  • Cryptanthla leiocarpa — Springer 1935 “the more open places in chaparral”
  • Pectocarya penicillata — Dengler [Notebook] 4/21/1963: “No Pectocarya penicillata-- could be found W. of Mimulus gully in chap. as last year (Rare)”
  • Phacelia californica — Dengler 1975; not reported since
  • Phacelia malvifolia — Dengler 1975; not reported since

BRASSICACEAE

  • Sinapis arvensis — Moeur 1947; not reported since
  • Turritis glabra (Arabis glabra var. g.) — Dengler 1975; not reported since

CAPRIFOLIACEA

  • Lonicera interrupta — Macklin 1988. Perhaps the same cream-flowered bush on Trail 2 extension SE of the brick cistern now referred to L. hispdula

CHENOPODIACEAE

  • Chenopodium murale — Dengler 1975; not reported since
  • Dysphania botrys (Chenopodium b.) — Dengler 1975; not reported since

EUPHORBIACEA

  • Euphorbia crenulata —  Dengler 1975; not reported since

FABACEA

  • Acacia baileyana — Dengler 1975; not reported since
  • Lathyrus tingitanus — Dengler 1975; not reported since
  • Lupinus microcarpus var. densiflorus X L. nanus — Dengler [Notebook] 4/14/63. See note
  • Trifolium wormskioldii — Moeur 1947; not reported since
  • Vicia ludoviciana ssp. ludoviciana — Porter 1962; not reported since

LAMIACEAE

  • Mentha arvensis — Springer 1935 “frequent in moist places”; Dengler 1975; not reported since
  • Prunella vulgaris var. vulgaris — Dengler 1975; not reported since

LYTHRACEAE

  • Lythrum californicum — Dengler 1975; not reported since

MALVACEAE

  • Malva parviflora — Dengler 1975; not reported since

OLEACEAE

  • Ligustrum ovalifolium — Dengler 1975; not reported since. May be same plant we call L. vulgare

ONAGRACEAE

  • Epilobium campestre (E. pygmaeum) — Seed hair-tuft 0. Dengler 1975; not reported since
  • Epilobium ciliatuml ssp. watsonii — Dengler 1975.
  • Ludwigia palustris — John Thomas 1979, Aquatic Plants Notes "John Thomas suggests the following as a very rough list of plants to be found in Searsville Lake"

OROBANCHACEAE

  • Castilleja attenuata X Castilleja exserta ssp. exserta — Dengler 1975; not reported since
  • Castilleja exserta  X  Castilleja rubicundula ssp. lithospermoides — Dengler 1975; not reported since, hybrid not mentioned in TJM2.
  • Castilleja subinclusa ssp.  franciscana — Dengler 1975; not reported since
  • Triphysaria versicolor ssp. versicolor — Springer 1935 and Moeur 1947; not reported since

OXALIDACEAE

  • Oxalis articulata ssp. rubra (O. rubra) — Dengler 1975; not reported since

POLEMONIACEAE

  • Leptosiphon ambiguous (Linanthus a.) — Moeur 1947; not reported since. Likely misidentification of Leptosiphon parviflorus

POLYGONACEAE

  • Rumex fueginus (R. maritimus) — Dengler 8/18/1963: “R. fueginus -- green, full achenes by laurel nearest C.M.Cr. near So corner of line - to 40" tangled in other growth. Inner parianth segments almost fringe”. Not reported since
  • Rumex transitorius (R. salicifolius var. transitorius) — Dengler 1975; not reported since

RANUNCULACEAE

  • Ranunculus aquatilis var. diffuses (R. a. var. capillaceus) — Dengler 1975; not reported since

RHAMNACEAE

  • Ceanothus thyrsiflorus — Dengler 1975; Dengler 1984 "Correction--deleted"

ROSACEAE

  • Poterium sanguisorba — Dengler 1975; not reported since

RUBIACEAE

  • Galium trifidum ssp. columbianum (G. t. var. pacificum) — Dengler 1975; not reported since

SALICACEAE

  • Salix sitchensis (S. coulteri) — Dengler 3/16/1964, See note

SCROPHULARIACEAE

  • Buddleja davidii — Dengler 1975; not reported since
  • Limosella aquatic — Dengler 1975; not reported since

SOLANACEAE

  • Solanum nigrum — Moeur 1947 “Rare, one plant in dry bed of SF Creek”

VALERIANACEAE

  • Plectritis congesta — Corolla 4–9.5 mm, pale to dark pink; spur tip generally enlarged; spur generally < 1/2 corolla tube length; fls fragrant. Dengler 1975; not reported since

ALLIACEAE

  • Allium serra — Springer 1935: “Infrequent in May in an openly wooded strip of serpentine”; D. Porter 1962: “serpentine soil of meadow and adjacent slopes.” Not reported since. Probably plants now called Allium peninsulare var. franciscanum

ARACEAE

  • Lemna gibba — Dengler 1975; not reported since

CYPERACEAE

  • Bolboschoenus robustus (Scirpus r.) — Dengler 8/26/1981 “SF Cr bed at Dennis Martin Settlement site”; not reported since
  • Carex senta — Dengler 1975; not reported since. CCH record for SF Cr. watershed
  • Cyperus esculentus — Dengler 1975; not reported since
  • Eleocharis acicularis var. acicularis — Dengler 1975; not reported since

HYDROCHARITACEAE

  • Elodea canadensis — Dengler 1975; not reported since

IRIDACEAE

  • Iris macrosiphon — Springer 1935, "April 1934 at edge of road through chaparral above lake"; no reported since

LILIACEA

  • Clintonia andrewsiana 
  • Prosartes smithii (Disporum s.) —

POACEAE

  • Phalaris minor — Dengler 1975; not reported since

THEMIDACEA

  • Muilla maritima — R. Porter no date or location; J. Funk 2011 in serpentine prairie (JRBP Brown Bag video 5/1/12). Probably Triteleia hyacinthina

ZANNICHELLIACEAE

  • Zannichellia palustris — 1979 mention by John Thomas as possibly to be found in lake in Aquatic Plant Notes

 

 

Table 2. Redeterminations and erroneous inclusions

ASTERACEAE
  • Corethrogyne filaginifolia — JROH1032, 1033 now ID Lessingia hololeuca.
  • Lessingia tenuis — Vouchers now ID L. pectinata Greene var. tenuipes (J. T. Howell) Markos. See note below.

BORAGINACEAE

  • Cryptantha muricata — Dengler 1975; not reported since. Misidentification of C. micromeres, collected on Westridge outside the Preserve in Wiliam Cooper's research plots.

BRASSICACEAE

  • Barbarea vulgaris — JROH1447 now ID B. orthoceras.
  • Descurainia sophia — JROH1506 collected on University lands outside JRBP.

CARYOPHYLLACEAE

  • Polycarpon depressum — JROH1710 now ID P. tetraphyllum var. t.; poor specimen.

CONVOLVULACEAE

  • Calystegia occidentalis ssp. o. — JROH1839, 1840, 1841 now ID Calystegia purpurata ssp. purpurata.

ERICACEAE

  • Arctostaphylos crustacea ssp. crinita — JROH1923, 1924, 1925, 1926 now ID A. c. ssp. crustacea.
  • Arctostaphylos tomentosa (Pursh) Abrams — JROH1922  annotated by Alice Eastwood as Arctostaphylos canescens Eastwood 

FABACEAE

  • Acacia decurrens — JROH1980 now ID A. dealbata.
  • Genista maderensis — JROH2006, 2007, 2008 now ID Genista monspessulana.

FAGACEAE

  • Quercus wislizeni var. wislizeni — JROH2527, 2528 now ID Q. wislizeni var. frutescens. See note below.

GENTIANACEAE

  • Centaurium davyi — Many vouchers now ID Zeltnera muehlenbergii; these sheets require further review.
  • Centaurium tenuiflorum  — JROH2579 now ID Zeltnera muehlenbergii.

MONTIACEAE

  • Claytonia parviflora ssp. viridis — JROH3035, 3336 now ID C. exigua ssp. exigua.

OROBANCHACEAE

  • Castilleja wightii — JROH4011 now ID Castilleja sp. See note below.
  • Orthocarpus purpurascens — Vouchers now ID Castilleja densiflora ssp. d. See note C. exerta.
  • Orobanche grayana ssp. jepsonii — JROH3086 now ID O. vallicola per Alison Colwell. See note below.

PLANTAGINACEAE

  • Collinsia franciscana (syn. Collinsia multicolor) —  Dengler 1963; not reported since. Mididentification of Collinsia sparsiflora.

POLEMONIACEAE

  • Gilia achilleifolia ssp. a. — Vouchers now ID G. achilleifolia ssp. multicaulis.

PRIMULACEAE

  • Primula clevelandii X P. hendersonii — JROH3384, 3386 now ID P. hendersonii. See note below.

URTICACEAE

  • Urtica urens — JROH4227 now ID Hesperocnide tenella.

ALLIACEAE

  • Allium dichlamydeum — Vouchers now ID A. peninsulare var. franciscanum. See note.

CYPERACEAE

  • Bolboschoenus robustus — JROH4421 now ID B. maritimus.

JUNCACEAE

  • Juncus mexicanus — JROH4475 now ID J. patens.

LILIACEAE

  • Calochortus venustus — Vouchers now ID Calochortus argillosus.

POACEAE

  • Bromus catharticus var. elatus — JROH4906, 4907 now ID Bromus catharticus var. catharticus.
  • Bromus pseudolaevipes— Vouchers now identified as B. carinatus (CCH  records for SMCo. & SCCo.)
  • Eragrostis diffusa —  JROH5140 now ID E. pectinacea
  • Eragrostis pilosa var. pilosa —  JROH5143 now ID E. pectinacea
  • Festuca occidentalis — JROH5715; See note.
  • Festuca pratensis — Vouchers now ID F. arundinacea.
  • Hordeum jubatum — JROH5121 now ID Elymus multisetus.
  • Melica geyeri — JROH5424 now ID Poa secunda ssp. s.
  • Poa unilateralis — JROH5267 now ID Koeleria macrantha.

THEMIDACEAE

  • Brodiaea coronaria — See note.
  • Muilla maritima — See note.

 

NOTES

Allium peninsulare var. franciscanum: reports of Allium Serra in 1935 and 1962 also likely A. p. var. f.

Apiastrum angustifolium: Dengler [Notebook] 10/27/1962 “SF Cr — beach 100' below falls cut by small rivulet last year. Growing in this shallow running water”; not reported since

Brodiaea coronaria var. coronaria is currently thought to have been misapplied locally to individuals of B. terrestris. However, additional examination of the matter is desirable; Dengler’s collection site (see below) has never been relocated and his voucher is missing. The Jepson eFlora (9/21//2016) defines the bioregional distribution of the taxa as Cascade Range Foothills, northern Sierra Nevada Foothills, and Sacramento Valley. The Jepson eFlora treatment also states:

Brodiaea coronaria subsp. coronaria has been misapplied to plants with tightly inrolled staminodes that are appressed to the stamens. At least 2 such groups are morphologically, ecologically distinguishable, one in montane meadows in KR, CaR, MP, southern Oregon, the other in grassland in ScV, northern SnJV. In addition, California plants morphologically similar to Brodiaea coronaria from the Pacific Northwest differ in chromosome number and may represent separate lineages of hybrID origin. Study needed.

A JROH sheet dated 05/16/1971 collected by Herb Dengler has been missing from sometime after 1977 when Ruth Porter annotated it. Location given as “Field S.W. of Searsville Rocks near largest stone in grassland.” CCH shows two other JRBP collections UCSB17659 (1963) and SEINET3884838 (1984), both reported from serpentine.

Castilleja exserta ssp. exserta is included on the plant list but lacks a voucher. Constructing a plausible just-so story explaining evIDence for it is problematic. Springer 1935 states “Abundant in April and May, often found with Orthocarpus densiflorus”; Dengler includes it on his 1975 plant list without comment along with the purported hybrid Castilleja exserta X Castilleja attenuata. One does occasionally find hybrid mini-swarms of Castilleja densiflora ssp. densiflora X Castilleja attenuata JROH5780, 4/26/2016). Specimen JROH3983, 4/3/1983, has Orthocarpus purpurascens on the original label but the sheet has been annotated Castilleja densiflora ssp. densiflora , a determination with which we agree in that the corolla beaks are straight, puberulent; filaments glabrous. The herbarium staff has searched carefully without success for 15 years for

Castilleja wightii: Poor specimen; widely-spaced cauline lvs and no branching proximal to infl.; herbage does appear glandular.

Festuca occidentalis: JROH5715 collected in 1903; not reported since. Specimen location dubious, however, Western fescue currently grows nearby in openings in woodland on the eastern slopes of the Santa Cruz Mountains at slightly higher elevations including along the Razorback RIDge Trail in the upper reaches of the Corte Madera Creek watershed and at mid-elevations in Huddart County Park and the Phleger Addition to GGNRA in the Union Creek drainage.

Lessingia tenuisLessingia glandulifera var. glandulifera: yellow-fl Lessingia sheets need additional review. After TJM2, some sheets are determined L. pectinata Greene var. tenuipes (J. T. Howell). Markos. "Fls yellow blooming May-Oct; specimens examined have purple banding near the throat of corolla tube and short triangular style appendage" (John Rawlings, 2017.

Lupinus microcarpus var. densiflorus L. nanus: Dengler [Notebook] 4/14/63 “Lupinus nanus x L. densiflorus–upper C.M.F.in bloom on hip of hill above fence particularly 50 ft. up. These L. nanus seem introgressed, but pods must be gathered Leaflets too broad for L. nanus and flowers more verticillate. Pods beginning, but already secund'”

Orobanche determinations problematic for specimens without indication of host; other characters such as corolla color and habitat also routinely lacking.

Quercus wislizeni var. wislizeni: Our earliest vouchers are identified Quercus wislizeni and precede publication of the name Quercus wislizeni var. frutescens. Ann Lambrecht reported Dengler had referenced in the 1970s a single example of Quercus wislizeni var. wislizeni in Sector 31 east of Rd E and Trail 12, south of Trail 11 in open woodland. However, a Dengler note dated 1993 in his annotated copy of Flora of the Santa Cruz Mountains indicates by that time he consIDered only var. frutescens present on the Preserve.

Perideridia gairdneri ssp. gairdneri : “There's only my memory of Herb taking several of us far off trail to show us this plant. Don't think that's good enough to be considered a second report" (A. Lambrecht). No CCH records for San Mateo Co. east of the Santa Cruz Mtns.

Primula clevelandii X P. hendersonii hybrids have been reported by Herb Dengler in 1959 and 1973 (JROH3384, JROH3386 respectively). Individuals not exactly matching taxa described in different manuals (Munz, Thomas, TJM) have been referred to P. hendersonii following TJM2. Dodecatheon confusion is not unique to our locality.

Salix sitchensis (S. coulteri):  first and only known report Dengler 3/16/1964: "Sausal Cr. (Couldn't find in Corte Madera Woodland where expected). With staminate flowers available." Since Dengler found staminate plant or plants its single stamen should have confirmed identification along with silky abaxial lf surface. Further, early collections from Portola are in CCH; all San Mateo Co, records. Unfortunately for our benefit Dengler failed to make a collection, note abundance, associates, and especially note the identifying characters. 

Zeltnera: Zeltnera davyi is documented for Stanford’s Academic Preserve, west of the Preserve. Centaurium sheets need additional review.

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